Legal Question in Family Law in California

I hate to give the typical lawyer answer, but "it depends." Only one county should have jurisdiction over the children at a time. So there should not be conflicting orders. Normally, there should not have been a guardianship order in a different county than the one that issued the custody orders. How and why that came to happen would probably reveal the answer to your question, but maybe not even then. Bottom line: that's a very specific and unusual situation that you can't really get an answer to on an internet Q&A service.